American politics uses "the wilderness" as a Biblical reference. Someone who is in the wilderness is an outcast, a nomad, someone without a seat in a place of power.
In practice, it is used to mean a politician or party lacking the power or influence normally due.
Example: "The Senator spent several years in the political wilderness after having made comments that offended members of a major religion. Only recently has he come back in favor."
In this sense, someone who is in the wilderness is like someone who is in the dog house.
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